For centuries, the Scythians ruled the vast Eurasian steppe from the Black Sea to the Altai Mountains. Described by Herodotus as fierce and mobile horsemen, they left behind kurgans—impressive burial mounds filled with weapons, gold, and sometimes mummified bodies. But who were the Scythians, genetically speaking? Where did they come from? And were they as homogeneous as classical authors imagined?
Thanks to ancient DNA research, their genetic story is now unfolding—and it reveals a dynamic, diverse, and mobile people shaped by multiple waves of migration across the steppes.
Genetic Clues from Three Key Studies
- Gnecchi-Ruscone et al. 2021 (Science Advances): Showed that Iron Age individuals from Kyrgyzstan (Alay Valley) carry mixed Steppe and Iranian-related ancestry.
Read the study - Matisoo-Smith et al. 2022 (Preprint): Focused on western Kazakhstan, revealing blends of Steppe, BMAC, and East Asian ancestry.
Read the study - Unterländer et al. 2019 (Current Biology): Demonstrated that Eastern and Western Scythians had different genetic backgrounds.
Read the study
A Mosaic of Origins
The term Scythian covered a wide range of Iron Age steppe cultures. Genetically, they descend primarily from Bronze Age Steppe herders, like the Yamnaya and Andronovo, but also show significant regional admixture. Western Scythians near Ukraine carry more European Neolithic ancestry, while eastern groups (e.g., Saka, Sarmatians, Tashtyk) reflect increasing Iranian and East Asian influence.

Genetic Comparison: East vs. West
To illustrate the diversity among these groups, we compared the genome-wide data (G25 coordinates) of five representative individuals:
| Sample ID | Culture | Region | Approx. Date | Cosine Distance to KE9609 (Eastern Tashtyk) |
|---|---|---|---|---|
| KE9609 | Tashtyk | South Siberia | ~200 CE | 0.000 |
| LC8544 | Tashtyk | South Siberia | ~200 CE | 0.023 |
| MJ-09 | Catacomb | Ukraine | ~2500 BCE | 0.201 |
| MJ-12 | Cimmerian | Ukraine | ~800 BCE | 0.577 |
| MJ-13 | Western Scythian | Ukraine | ~600 BCE | 0.322 |
As the distance values show, KE9609 and LC8544 are genetically very close—both belong to the Siberian Tashtyk horizon. In contrast, Western Scythians like MJ-13 are more distant, and Cimmerians like MJ-12 are even more distinct, despite sharing similar nomadic lifeways.
The Scythian Legacy
Rather than being a single people, the Scythians represent a **cultural network** of horse-riding elites who dominated the steppe through trade, mobility, and warfare. Their genetic diversity tells the story of migrations, local integration, and continental-scale contact zones between Europe and Asia.
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Kazakhstan_Berel_IA:BRE010,0.094473,-0.116786,0.050157,0.04199,-0.048624,-0.005299,0.009165,0.006231,-0.01493,-0.016948,-0.007632,-0.001199,-0.006838,-0.021882,0.012486,0.009281,-0.013299,0.001267,0.00176,0.005002,-0.015972,0.000371,-0.0053,0.001084,-0.000359 Kazakhstan_Berel_IA:BRE011,0.064879,-0.173656,0.062979,0.033269,-0.05201,-0.008646,0.00517,0.012923,-0.010431,-0.02041,-0.016239,0.00045,0.00773,-0.017478,0.014658,-0.005038,-0.01708,-0.001014,0.00352,-0.001501,-0.01722,0.003091,-0.012448,-0.000482,0.007305 Kazakhstan_Berel_IA:BRE012,0.087644,-0.088351,0.060339,0.055233,-0.054164,0.006972,-0.00329,-0.003923,-0.014317,-0.034443,-0.008282,0.000749,0.002973,-0.027249,0.008143,0.002784,-0.01343,-0.00266,0.006536,0.006378,-0.030321,-0.00136,0.003451,-0.007471,-0.005628 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Kazakhstan_Sarmatian_IA:CLK001,0.097888,0.054839,0.037712,0.076551,-0.029236,0.020359,0.0047,-0.010846,-0.02986,-0.045559,0.012342,0.005245,0.001338,-0.022708,0.0076,0.023866,-0.011735,-0.009502,0.009553,-0.007253,-0.013102,-0.008408,-0.011955,0.010845,-0.006466 Kazakhstan_Saka_IA:CSP002,0.081953,-0.098506,0.059208,0.05814,-0.036314,0.00502,-0.00376,-0.000923,-0.011044,-0.006925,-0.013803,0.007343,-0.000446,-0.023258,0.020358,0.008618,0.005998,-0.012669,-0.001257,0.011005,-0.016097,-0.005441,0.001109,-0.006386,-0.001317 Kazakhstan_Saka_IA:CSP002,0.081953,-0.098506,0.059208,0.05814,-0.036314,0.00502,-0.00376,-0.000923,-0.011044,-0.006925,-0.013803,0.007343,-0.000446,-0.023258,0.020358,0.008618,0.005998,-0.012669,-0.001257,0.011005,-0.016097,-0.005441,0.001109,-0.006386,-0.001317 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Kazakhstan_Birlik_Tasmola_EIA:BIR013,0.034147,-0.372699,0.065619,-0.023256,-0.05601,-0.03765,0.016921,0.019615,0.010022,0.005832,-0.029879,-0.003747,0.001338,-0.005643,-0.003122,-0.003713,0.003912,0.002154,0.000754,0.015007,-0.022585,-0.009274,-0.016022,0.001446,0.001197 Kazakhstan_Birlik_IA:BIR010,0.052359,-0.295519,0.069767,0.005491,-0.072629,-0.031236,0.016216,0.020076,-0.000409,-0.006196,-0.021922,-0.0003,0.003122,-0.013349,0.004614,-0.005569,-0.012517,0.002027,0.007165,0.012006,-0.017344,0.003091,-0.01023,-0.004338,-0.002754 Kazakhstan_Korgantas_IA:KBO001,0.043253,-0.386917,0.088623,-0.015181,-0.077245,-0.04769,0.007285,0.02723,0.010022,0.006196,-0.018837,0.000899,-0.006392,0.000826,-0.000271,0.002254,-0.004433,-0.003674,0.001508,0.011631,-0.021712,0.001484,-0.01479,-0.003856,0.000958 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Kyrgyzstan_Saka_IA:KEN001,0.095611,-0.022342,0.022627,0.050065,-0.041546,0.014223,0.00047,0.003231,-0.02352,-0.028429,-0.008282,0.00045,0.006392,-0.01968,0.023072,0.009546,-0.007432,-0.004814,-0.005782,0,-0.007237,-0.000247,-0.000739,0.003253,0.002874 Kyrgyzstan_Saka_IA:KEN002,0.086506,-0.036559,0.02753,0.05814,-0.040315,-0.001394,-0.00752,0.005307,-0.02168,-0.03426,-0.002436,0.00015,0.001933,-0.007569,0.023887,0.010209,-0.008475,0.004181,0.003645,-0.000625,-0.007487,0.002597,0.002711,-0.00012,-0.000359 Kyrgyzstan_Saka_IA:KEN003,0.099026,-0.025388,0.028284,0.05814,-0.025851,0.021753,0.00329,-0.007846,-0.02168,-0.036447,-0.010068,-0.011989,0.012042,-0.016239,0.031623,0.004906,-0.01708,0.004181,-0.003897,0.000875,-0.016845,-0.002968,-0.002835,-0.004338,-0.004071 Kazakhstan_Sarmatian_Turk_IA:KKB001,0.066017,-0.168578,0.049403,0.022287,-0.047086,-0.006972,0.005405,0.005769,-0.007772,-0.009841,-0.01494,-0.005245,0.000149,-0.016102,0.010043,0.012198,0.005215,0.002407,0.003771,0.005878,-0.013601,-0.001113,-0.008134,0.001566,0.001437 Kazakhstan_Sarmatian_Turk_IA:KKB002,0.079676,-0.175687,0.060339,0.013243,-0.050471,-0.011156,0.000705,0.004154,-0.01268,-0.005103,-0.013153,-0.00045,-0.003122,-0.017478,0.007872,0.010475,0.007041,0.002534,0.001885,0.010255,-0.022585,0.009274,-0.00912,0.010122,-0.005269 Kazakhstan_Tasmola_EIA:KKM001,0.084229,-0.113739,0.057699,0.049096,-0.033852,-0.001673,-0.00188,0.003231,-0.006136,-0.012028,-0.01153,0.004946,0.009514,-0.025323,0.014386,0.010607,-0.00665,0.006081,-0.002263,0.005002,-0.011605,0.00272,0.001109,0.001928,-0.000359 Kazakhstan_Kayalyk_Medieval:KLK002,0.035285,-0.385901,0.090509,-0.018734,-0.07663,-0.050479,0.022561,0.021922,0.00225,0.017312,-0.029879,0.003447,-0.000743,0.005367,0.011401,0.001591,-0.01004,-0.00038,0.013073,0.027763,-0.023334,-0.017682,-0.033647,0.001325,-0.002275 Kazakhstan_OtrarCulture_LIA:KNT001,0.09675,0.063978,-0.013199,0.0323,-0.036314,0.016455,0.002585,-0.003923,-0.033542,-0.029887,-0.005846,-0.005695,0.011447,-0.023121,0.018458,0.023866,0.004824,-0.002534,-0.001508,0.000375,0.002745,-0.002102,-0.004067,0.010724,0.002874 Kazakhstan_OtrarCulture_LIA:KNT002,0.10927,0.060932,0.009805,0.048127,-0.028621,0.023148,0.00517,-0.007615,-0.024952,-0.031345,0.006983,-0.003897,-0.002825,-0.013625,0.019001,0.015248,-0.01004,-0.001774,0.002765,-0.005503,-0.017344,-0.002102,-0.000493,0.002048,0.001557 Kazakhstan_OtrarCulture_LIA:KNT003,0.100164,0.059916,-0.006411,0.038437,-0.035391,0.018965,-0.00047,-0.014769,-0.037019,-0.028793,-0.000974,-0.001948,0.005946,-0.024772,0.020629,0.022142,-0.000261,-0.007348,-0.004651,-0.002501,-0.006114,-0.005317,-0.002342,-0.006989,0.004311 Kazakhstan_OtrarCulture_LIA_o:KNT004,0.067156,-0.136081,0.000754,-0.020026,-0.039392,-0.006136,0.012456,0.012692,-0.01084,-0.00328,-0.015265,-0.005995,0.007582,-0.004542,0.001493,0.005171,-0.006519,-0.000633,0.003771,-0.004002,-0.011605,-0.004451,-0.003204,-0.004579,-0.00012 Kazakhstan_OtrarCulture_LIA_o:KNT005,0.084229,0.071087,-0.072407,0.04199,-0.066782,0.026774,0.006815,0.002538,-0.027611,-0.024055,-0.003085,-0.004496,-0.001338,-0.007569,0.0076,0.017369,-0.007171,0.00076,0.005405,-0.02101,-0.00262,0.000247,0.003328,-0.003615,0.006826 Kazakhstan_Hun_Elite_LIA:KRY001,0.056912,-0.30669,0.069767,-0.011305,-0.05878,-0.018965,-0.000235,0.001615,0.00634,-0.005467,-0.031503,-0.008692,-0.003419,0.001101,0.008143,-0.004508,-0.008475,-0.003167,-0.000754,0.014382,-0.028699,0.001113,-0.008134,0.00253,0.005029 Kazakhstan_Tasmola_EIA:KSH001,0.080814,-0.119832,0.060716,0.046189,-0.046162,0.001952,0.00188,-0.008077,-0.014317,-0.025878,-0.01153,-0.002248,0.002081,-0.023946,0.015879,0.012463,0.004433,0.003927,-0.003142,0.004752,-0.024582,0.004204,0.00456,0.002771,-0.002754 Kazakhstan_Tasmola_EIA:KSH002,0.083091,-0.112724,0.058077,0.046835,-0.038161,0.011435,0.008225,0.006692,-0.011044,-0.020957,-0.012342,-0.005245,0.0055,-0.014175,0.007193,0.003447,-0.001173,-0.003927,-0.003645,0.005503,-0.01697,0.004575,0.00037,0.00241,0.005868 Kazakhstan_Tasmola_IA:KSH003,0.084229,-0.11577,0.05506,0.039083,-0.048932,0.00502,-0.00235,-0.001154,-0.013703,-0.016037,-0.01429,-0.010191,0.007582,-0.026974,0.017372,0.008884,0.003781,-0.00266,-0.00088,0.01013,-0.01672,0.001978,-0.001849,0.002651,0.002634 Kazakhstan_Tasmola_EIA:KSH004,0.0774,-0.153345,0.055437,0.033915,-0.045239,0.00251,0.00282,0.000462,0.000614,-0.017859,-0.011367,-0.004946,0.002081,-0.016928,0.009365,0.009812,0.010952,-0.00038,-0.004651,0.004752,-0.017344,0.003215,-0.008751,0.002048,-0.006945 Kazakhstan_Sarmatian_IA_o1:KSK002,0.10927,0.073118,0.038843,0.066538,-0.011387,0.029562,0.002585,0.006461,-0.022907,-0.028429,-0.008607,0.005845,0.003568,-0.015551,0.032844,0.007292,-0.032857,0.010135,0.008045,-0.005753,-0.011729,0.004946,0.002958,-0.002892,0.000359 Kazakhstan_Tasmola_EIA:KYZ001,0.079676,-0.120848,0.058077,0.047481,-0.035699,-0.002789,0.00047,0.002769,-0.013908,-0.019499,-0.003085,-0.003597,0,-0.018992,0.013979,0.01127,0.00326,-0.006334,-0.005908,0.005878,-0.021213,0.005935,-0.007272,0.007471,-0.000239 Kazakhstan_Tasmola_EIA:KYZ002,0.076261,-0.152329,0.062225,0.042636,-0.040623,-0.001952,0.003525,0.003923,-0.010226,-0.01549,-0.019974,-0.002847,0.008176,-0.017065,0.004614,-0.002519,-0.016037,-0.004434,-0.00817,0.003752,-0.021961,0.009892,-0.004067,-0.003133,-0.000718 Kazakhstan_Tasmola_EIA:KZL001,0.093335,-0.093429,0.059208,0.054587,-0.041238,0.017849,0.00188,-0.005307,-0.017794,-0.031162,-0.005521,-0.001349,-0.003717,-0.01913,0.024837,-0.002121,-0.016689,0.00152,0.000251,0.001751,-0.01148,0.006059,-0.002218,0.001928,-0.000838 Kazakhstan_Tasmola_LBA:KZL004,0.120652,0.105615,0.062602,0.08398,0,0.038766,0.00658,0.000923,-0.013703,-0.034989,-0.000487,0.000599,-0.000595,-0.02257,0.019272,0.014585,-0.008084,-0.004814,-0.005405,0.002126,0.002371,-0.001484,0.002711,-0.001084,-0.001317 Kazakhstan_TurkicPossible_EIA:MMR001,0.069432,-0.189904,0.032809,0.007106,-0.044008,-0.004183,0.006815,0.01823,-0.011862,-0.010023,-0.021273,-0.002698,0.003271,-0.009358,0.007872,-0.003447,-0.012256,-0.003801,0.00176,0.002126,-0.012478,0.000371,0.005793,-0.005302,-0.010777 Kazakhstan_Tasmola_IA:NUR002,0.084229,-0.148267,0.069013,0.047481,-0.03416,0.007251,0.00094,-0.006,-0.010226,-0.022415,-0.009906,-0.003447,0.004014,-0.01445,0.013436,-0.003845,-0.013951,-0.00152,-0.004777,0.002876,-0.020464,0.00136,-0.003451,-0.002651,-0.010179 Kazakhstan_Sarmatian_IA_o2:SBL001,0.081953,-0.128972,0.055814,0.043605,-0.036007,0.010319,0.0047,0.003461,-0.011249,-0.024784,-0.009906,0.003297,0.002973,-0.020919,0.016694,0.007558,0,0.012415,0.007416,0.012756,-0.02009,-0.000371,-0.003451,0.01928,0.000599 Kazakhstan_Sarmatian_IA:SGZ001,0.114961,0.045699,0.035072,0.063954,-0.025235,0.020359,0.000235,0.002077,-0.023725,-0.038087,-0.005684,0.002997,-0.003122,-0.028213,0.02633,0.019225,0.009909,-0.002787,-0.000503,0.001876,-0.010357,0.005935,0.003081,0.021087,-0.010538 Kazakhstan_Sarmatian_IA:SGZ002,0.106994,0.04773,0.03017,0.07106,-0.032621,0.028726,0.001645,0.000923,-0.024747,-0.038816,-0.005521,0.003147,-0.001041,-0.005505,0.025651,-0.000133,-0.023991,0.003674,0.000251,-0.003001,-0.012104,0.005935,0.002958,0.001446,0.002634 Russia_EIA_Sargatka_IA:SHD001,0.095611,-0.067025,0.068259,0.058786,-0.047393,-0.00251,0.00846,0.003923,-0.010226,-0.028064,0.012666,-0.006744,0.009366,-0.031378,0.007736,-0.00305,-0.012647,-0.001774,-0.005154,-0.001751,-0.013975,0.01051,-0.001356,-0.005422,0.011616 Russia_EIA_Sargatka_IA:SMV001,0.087644,-0.102568,0.073161,0.055879,-0.056318,-0.009482,-0.001645,-0.002308,-0.018203,-0.036265,0.018999,-0.006894,0,-0.041837,-0.000271,0.004906,0.000652,0.003294,-0.003394,-0.002876,-0.004243,0.013231,0.006902,0.010845,-0.005269 Russia_EIA_Sargatka_IA:SMV002,0.093335,-0.06804,0.055814,0.059755,-0.053856,0.003347,0.004465,0.006923,-0.014726,-0.027335,0.009743,0.001199,0.00892,-0.032892,-0.000136,0.01074,0.000261,0.001014,-0.003645,-0.008129,-0.003119,0.009645,0.004314,0.004579,-0.000838 Kazakhstan_Tasmola_EIA:SRK001,0.0774,-0.15436,0.056191,0.034561,-0.035699,-0.00753,0.006345,0.008538,-0.009408,-0.008201,-0.013641,-0.003897,-0.002973,-0.020781,0.013301,0.009414,0.000782,-0.003294,0.003645,0.004252,-0.015223,0.002844,-0.000123,-0.003856,0.003233 Kazakhstan_Tasmola_EIA:TAL003,0.088782,-0.090382,0.064865,0.054587,-0.039084,0.008088,-0.001645,-0.007384,-0.014521,-0.022597,-0.009906,-0.005095,0.008474,-0.016652,0.020901,0.023071,0.004172,0.001394,0.000628,0.007003,-0.01984,0.005935,0.00037,0.006266,0.000599 Kazakhstan_Tasmola_IA:TAL004,0.08992,-0.109677,0.058077,0.053941,-0.042469,0.004183,-0.000235,0.001385,-0.010431,-0.027335,-0.007307,0.001349,0.006987,-0.024359,0.014658,0.010872,-0.002608,0.004054,-0.005782,0.009129,-0.011979,0.005935,0.001849,-0.003374,0.002994 Kazakhstan_Tasmola_EIA:TAL005,0.091058,-0.107646,0.056568,0.04522,-0.024312,0.015618,-0.002585,0.003923,-0.017385,-0.013485,-0.013478,0.00015,0.008028,-0.026836,0.011672,0.003978,0.000391,-0.005574,-0.002263,0.004877,-0.019715,0.005564,0.003944,-0.003856,0.002155 Kazakhstan_Tasmola_EIA:WAR001,0.087644,-0.101553,0.049026,0.052003,-0.047086,-0.002789,0.00705,-0.003461,-0.011658,-0.031162,-0.010393,-0.003147,0.001635,-0.018579,0.022122,0.009679,-0.009648,-0.003294,-0.003897,-0.003627,-0.01697,0.00507,0.003451,0.003374,-0.004311
References
- Gnecchi-Ruscone, G. A., et al. (2021). Science Advances. https://doi.org/10.1126/sciadv.abe4414
- Matisoo-Smith, E., et al. (2022). Preprint. https://doi.org/10.21203/rs.3.rs-1993191/v1
- Unterländer, M., et al. (2019). Current Biology. https://doi.org/10.1016/j.cub.2019.06.019