The edges of the Roman Empire were never barren limits. They were dynamic zones of exchange, settlement, and transformation. Thanks to archaeogenetics, we now have the tools to explore these liminal spaces through the DNA of those who lived and died there. Three recent studies, from Valkenburg (South Holland), Sint-Truiden (Belgium), and a broad survey across Europe, offer new insights into the ancestry and mobility of Roman frontier populations. When combined with genetic data from Merovingian-era cemeteries, a fascinating story emerges: one of continuity, integration, and transition.

I. Valkenburg: A Cosmopolitan Outpost

The Roman military cemetery at Valkenburg, explored in the 2024 study by van Duijvenbode et al. (bioRxiv), presents a diverse genetic landscape. Most males carried the Y-haplogroup R1b1a1b1a1a2, associated with Iron Age Western Europeans, suggesting local recruitment. Yet, the mitochondrial DNA (mtDNA) lineages were notably varied: U5, H1-H11, T2, J1c, K1a, N1a, and even L2b, which points to African ancestry. These maternal haplogroups likely represent women of diverse geographic origin, some possibly arriving via military or economic networks.

 

Sample ID Y-DNA mtDNA
CGG107735 R1b1a1b1a1a1c1a H6a1b4
CGG107736
HV0+195
CGG107742
H4a1a1a3
CGG107744
N1a1a1a2
CGG107745 R1b1a1b1a1a1c1a H6a1b4
CGG107746 R1b1a1b1a1a2c1a1k U5a1a1
CGG107747 R1b1a1b1a1a2 H1+16311
CGG107748 R1b1a1b1a1a2 J1c1
CGG107749
L2b
CGG107750 R1b1a1b1a1a2 T2b6a
CGG107751 R1b1a1b1a1a1c1a2b1 U5a1b1
CGG107752 R1b1a1b1a1a2b1a1~ H8c1
CGG107753 J2a1a1b2a1b1b~ K1a4
CGG107754 R1b1a1b1a1a1c1 J1c3g
CGG107755
H1ae
CGG107756 R1b1a1b1a1a2b1 T2c1d1
CGG107757 R1b1a1b1a1a1c1a1 T2b
CGG107758 R1b1a1b1a1a3a1 U5b2a2
CGG107759
I2
CGG107760
T2b6a
CGG107761 R1b1a1b1a1a2e1 K1a3a1
CGG107762 I1a1b1 I2a
CGG107764
HV16
CGG107766 R1b1a1b1a1a2b H11a
CGG107767 R1b1a1b1a1a1c H2a2b
CGG107768 I2a1b1a1b1a1a~ K1a3

Autosomal data reinforces this narrative of admixture: while many individuals showed Western European profiles, others carried signals from Southern Europe and the eastern Mediterranean. One remarkable case is individual CGG107753, buried at Valkenburg, whose genome reflects a strong affinity with Byzantine or eastern Mediterranean ancestry, highlighting the reach of Roman-era mobility. Strontium isotope analysis confirmed high mobility, especially among non-local women.

II. Sint-Truiden: Diversity Within the Ranks

In 2025, Cassidy et al. (Genome Biology) revealed the genetic profiles of 42 individuals from Sint-Truiden, a Roman military site in Belgium. Here, the genetic diversity is even more striking: individuals trace ancestry to the Balkans, the Levant, North Africa, and beyond. Over one-third were born outside the region, as shown by isotopic data.

This pattern reflects the Roman military's recruitment from across the Empire and the permeability of provincial society. Like Valkenburg, Sint-Truiden supports the idea of rapid social integration: first-generation migrants intermarried, and their descendants formed a new, local population.

 

IV. Merovingians: Inheriting the Empire’s Diversity?

What happened after Rome? Genetic data from over 40 Merovingian individuals (KOS, OLV, WPK), as described in the study Capturing the fusion of two ancestries and kinship structures in Merovingian Flanders, offers a glimpse. Individuals like KOS009, OLV072, and OLV065 illustrate the persistence of diverse ancestries in early medieval Flanders, with varying degrees of admixture reflecting both continuity from the Roman period and new kinship structures emerging in Merovingian society. Many carried R1b male lineages and autosomal profiles similar to local Roman-period samples. Others, however, diverged, suggesting heterogeneity in post-Roman population structures.

Compared to the Valkenburg and Sint-Truiden individuals, the Merovingians seem genetically closer to local Romanized populations than to the more diverse military recruits. Some admixture components, possibly from earlier Roman migrations, persisted. This suggests that while the Roman Empire collapsed politically, its genetic legacy endured.

V. Conclusion: The Genetic Echoes of Empire

The Roman frontier was more than a military boundary, it was a corridor of human movement and exchange. From Valkenburg’s cosmopolitan cemetery to Sint-Truiden’s diverse garrison, and into the Frankish age, DNA tells a story of integration and inheritance.

The peoples of early medieval Europe did not arise from isolation but from centuries of contact. The Empire may have fallen, but its biological impact shaped the genetic landscape of its former provinces for generations to come.

 

For those interested in further analysis or visualization, here are the G25 coordinates of the samples:

G25 · V. Conclusion: The Genetic Echoes of Empire
CGG107735_Netherlands_South-Holland_Roman_IA,0.125205,0.128972,0.055814,0.055556,0.030467,0.024263,0.00376,-0.000692,0.000205,-0.00656,-0.00065,-0.004646,-0.014569,-0.011147,0.012893,0.008751,-0.012126,-0.001267,-0.000754,0.01063,-0.006364,0.002597,0.005793,0.015303,0.000958 CGG107736_Netherlands_South-Holland_Roman_IA,0.124067,0.139128,0.070899,0.043928,0.018465,0.0251,-0.010575,0.006923,-0.002454,-0.004191,0.002598,-0.009591,0.004906,-0.001514,0.029858,0.008088,-0.026859,-0.000253,0.006411,0.022136,0.004991,-0.005317,-0.008134,0.014821,-0.013891 CGG107742_Netherlands_South-Holland_Roman_IA,0.126344,0.144205,0.064488,0.060724,0.016003,0.017849,0.00329,-0.009461,-0.006954,-0.019317,-0.002598,0.001049,-0.0055,-0.022432,0.029994,0.008221,0.007562,0.005574,-0.004777,0.012381,0.001373,0.002844,-0.001109,0.017472,0.003233 CGG107744_Netherlands_South-Holland_Roman_IA,0.132035,0.139128,0.061848,0.052649,0.042777,0.021753,0.015746,0.006692,0.009204,-0.000547,-0.000487,0.004796,-0.010555,-0.019405,0.020087,0.015115,0.008345,-0.001267,0.002263,0.014882,0.004118,-0.000989,0.001972,0.006868,0.00455 CGG107745_Netherlands_South-Holland_Roman_IA,0.121791,0.123895,0.061471,0.059109,0.032929,0.028168,-0.002585,0.001846,0,-0.008565,-0.000812,-0.001199,-0.011893,-0.01734,0.018865,-0.001724,-0.0103,-0.003421,-0.002514,0.012256,0.006489,0.001484,0.001849,0.01928,-0.004071 CGG107746_Netherlands_South-Holland_Roman_IA,0.12862,0.146236,0.061471,0.031008,0.048009,0.016176,0.002585,0.005307,0.012271,0.020046,-0.001461,0.016935,-0.017393,-0.005367,0.0095,-0.009414,-0.019427,0.00266,0.005154,-0.005503,0.010981,-0.002102,-0.003574,-0.000602,-0.002275 CGG107747_Netherlands_South-Holland_Roman_IA,0.124067,0.140143,0.048271,0.022287,0.0437,0.009482,-0.001175,0.004384,0.007567,0.019499,-0.006008,0.004796,-0.016799,-0.005918,0.008958,-0.002121,-0.005867,0.000253,0.00817,-0.003877,0.00262,-0.000989,-0.000616,0.00976,0.001796 CGG107748_Netherlands_South-Holland_Roman_IA,0.137726,0.129988,0.055437,0.062016,0.040931,0.010877,0,0.004154,0.006545,-0.002005,-0.005359,0.004946,-0.00996,-0.017753,0.025515,0.02201,0.005085,0.008488,0.002388,0.009129,0.009234,0.003586,0.004067,0.014942,-0.002754 CGG107749_Netherlands_South-Holland_Roman_IA,0.132035,0.129988,0.075047,0.062985,0.044316,0.026495,-0.001175,0.005769,-0.003272,-0.006743,-0.004222,-0.001798,-0.011596,-0.011973,0.019544,0.012331,-0.002086,0.002534,-0.000377,0.003752,0.012104,-0.00272,0.002835,0.016026,0.000479 CGG107750_Netherlands_South-Holland_Roman_IA,0.124067,0.140143,0.058831,0.031977,0.048624,0.01004,0.009165,0.007615,0.010226,0.007289,-0.005359,0.004196,-0.011596,-0.008808,0.013301,0.002917,-0.004694,0.001647,0.004525,0.007504,0.001872,0.006554,0.004314,-0.003976,-0.002155 CGG107751_Netherlands_South-Holland_Roman_IA,0.125205,0.129988,0.06939,0.061693,0.028313,0.023427,0.003055,0.005307,0.003272,-0.000911,-0.000487,0.005545,-0.013825,-0.013349,0.027823,-0.005171,-0.025295,0.004941,-0.003394,0.005503,-0.00025,0.008903,-0.004314,0.008917,0.003113 CGG107752_Netherlands_South-Holland_Roman_IA,0.133173,0.127957,0.052797,0.033915,0.044931,0.014781,0.002585,0.002538,0.012885,0.008383,-0.004384,0.005845,-0.013231,-0.025873,0.014522,0.008221,-0.005867,-0.003801,0.00176,0.009505,-0.000873,-0.003833,0.001725,0.005784,0.002515 CGG107753_Netherlands_South-Holland_Roman_IA,0.10927,0.151314,-0.033941,-0.075582,0.002154,-0.024542,0.00188,-0.00923,-0.009408,0.023326,-0.002436,0.003747,-0.013677,0.000413,-0.016829,-0.002254,0.009127,-0.000887,0.003394,-0.006378,-0.013975,-0.002968,0.000739,0.001446,0.000958 CGG107754_Netherlands_South-Holland_Roman_IA,0.127482,0.135065,0.064865,0.056848,0.043393,0.018965,0.012221,0.005307,0.003272,-0.002187,-0.004384,0.001499,-0.008176,-0.016515,0.015065,0.001989,-0.006258,0.002407,0.002891,0.005253,0.003743,0.011129,-0.000863,0.006868,-0.003353 CGG107755_Netherlands_South-Holland_Roman_IA,0.127482,0.139128,0.053551,0.032946,0.044931,0.00753,-0.003055,0.007846,0.00409,0.012574,-0.007795,0.01169,-0.017839,-0.014863,0.018729,0.001193,-0.00678,0.005194,-0.004525,0.001876,-0.002745,0.004328,-0.000616,0.009399,-0.002155 CGG107756_Netherlands_South-Holland_Roman_IA,0.12862,0.141159,0.048271,0.026163,0.040623,0.005857,0.00329,0.002538,0.012271,0.01877,-0.00682,-0.00015,-0.01888,-0.004679,0.013436,0.009546,-0.000391,-0.00266,0.000628,-0.001751,0.000499,0.009645,-0.001356,0.00253,0.004071 CGG107757_Netherlands_South-Holland_Roman_IA,0.125205,0.132019,0.075424,0.066538,0.040931,0.021475,0.000235,0.002538,0.009613,-0.005467,-0.005521,0.008542,-0.00996,-0.01156,0.021987,0.015911,0.001043,-0.002914,0.001131,0.014007,0.01123,0.006801,-0.001972,0.011447,0.006107 CGG107758_Netherlands_South-Holland_Roman_IA,0.129758,0.12491,0.059962,0.053295,0.034468,0.019522,0.006815,0.000923,0.001227,0.002551,-0.008931,0.003297,-0.008771,-0.009496,0.020358,0.019358,0.008084,-0.00114,0.006662,0.009505,0.00574,0.005441,-0.000986,0.00735,-0.001916 CGG107759_Netherlands_South-Holland_Roman_IA,0.124067,0.137096,0.056191,0.034561,0.037545,0.014781,-0.00235,0.007846,0.014521,0.013303,0.00065,0.004646,-0.015461,-0.016239,0.008415,0.002121,-0.00339,0.002787,0.003771,-0.001251,0.008111,0.011005,-0.001972,0.012893,-0.003592 CGG107760_Netherlands_South-Holland_Roman_IA,0.133173,0.139128,0.063356,0.053295,0.03693,0.021753,0,0.008077,-0.002045,-0.004556,-0.007145,0.002997,-0.012339,-0.013625,0.018865,-0.001326,-0.014864,0.002407,0.004525,-0.001251,0.003119,-0.002226,0.001479,0.004579,-0.000599 CGG107761_Netherlands_South-Holland_Roman_IA,0.140002,0.136081,0.06675,0.06783,0.045547,0.017291,0.001645,-0.000923,-0.001023,-0.002734,-0.002598,0.006594,-0.01219,-0.01679,0.016422,0.008088,0.000652,0.006334,0.006411,0.004127,0.00574,0.000989,-0.001972,0.008796,-0.003353 CGG107762_Netherlands_South-Holland_Roman_IA,0.129758,0.132019,0.09164,0.080104,0.046778,0.029841,0.00987,0.011307,-0.000614,-0.035354,-0.001949,-0.009142,0.016501,0.013487,-0.003122,0.01127,0.00665,0.009248,0.006788,0.01013,0.000873,-0.007172,0.001232,0.000482,0.00479 CGG107764_Netherlands_South-Holland_Roman_IA,0.122929,0.137096,0.058831,0.05814,0.041546,0.022869,-0.001175,0.002538,0.00634,0.002005,0.000162,0.008992,-0.015758,0.004404,0.019408,-0.010607,-0.021905,-0.000633,0.00264,0.002251,0.009109,0.001237,0.005546,0.008314,-0.003233 CGG107766_Netherlands_South-Holland_Roman_IA,0.130897,0.125926,0.076178,0.066861,0.033545,0.025937,0.012691,0.002769,-0.003272,-0.003462,-0.000162,0.004346,-0.008325,-0.011285,0.02443,-0.002917,-0.014603,0.002407,0.000126,-0.003752,0.012228,0.00915,-0.000493,0.013255,-0.004191 CGG107767_Netherlands_South-Holland_Roman_IA,0.12862,0.131003,0.058831,0.055879,0.032929,0.019243,0.008225,0.006923,-0.003272,0.001822,0.003735,0.004946,-0.006541,-0.013625,0.022801,0.009679,-0.009909,0.005954,0.000503,0.002501,0.001996,0.003215,0.002095,0.01217,0.00012 CGG107768_Netherlands_South-Holland_Roman_IA,0.129758,0.129988,0.061471,0.059755,0.031083,0.02761,0.002585,0.004846,0.000205,-0.007836,-0.003248,0.005695,-0.013528,-0.019955,0.025923,0.015513,-0.000652,-0.00114,-0.001131,0.004127,0.007487,-0.00136,-0.001972,0.010122,0.005269 Merovingian:KOS001,0.1161,0.141159,0.046009,0.036499,0.025851,-0.003904,0.006345,0.007846,0.009408,0.004374,0.000325,0.006594,-0.00446,-0.018304,0.015201,0.003447,-0.00339,0.002027,0.002765,0.000875,0.007986,0.011623,0.00419,0.013014,-0.006347 Merovingian:KOS002,0.119514,0.127957,0.058831,0.039083,0.031083,0.006414,0.0047,0.010615,0.009204,0.010752,-0.004547,0.003747,-0.013528,-0.010184,0.017508,0.021214,0.004694,0.006208,0.013073,0.001626,0.008485,-0.002597,-0.007395,0.010845,0.001557 Merovingian:KOS003,0.126344,0.132019,0.062979,0.0323,0.044316,0.015618,0.009165,0.005077,0.007363,0.020046,0.00065,0.009142,-0.012636,-0.008257,0.014522,0.008088,0.002217,0.003294,-0.004902,-0.003752,0.008235,-0.01286,-0.011955,0.007591,-0.002036 Merovingian:KOS006,0.120652,0.140143,0.064111,0.045866,0.035699,0.01506,0.00329,0.002538,0.010022,0.003645,-0.003573,0.005245,-0.010852,-0.013487,0.027551,0.012994,0.011735,-0.004434,0.005908,0.015132,0.004118,0.002473,0.001109,0.01205,-0.00467 Merovingian:KOS007,0.122929,0.125926,0.07203,0.059109,0.044008,0.021753,0.000235,0.001385,0.005113,-0.002734,-0.017376,3e-04,-0.015015,-0.012248,0.028908,0.005304,0.002086,0.002154,0.006285,-0.003252,0.009982,0.00272,-0.002218,0.015303,0.008502 Merovingian:KOS008C,0.122929,0.138112,0.058454,0.041021,0.041546,0.009203,-0.002585,0.002769,0.014726,0.009841,-0.002273,0.004946,-0.015907,-0.007569,0.030537,0.008221,-0.01017,-0.003927,0.004902,0.002376,0.004243,0.011994,0.000616,0.00976,-0.003712 Merovingian:KOS009,0.112685,0.142174,0.039975,0.017119,0.040623,0.007809,-0.003995,0.007846,0.017589,0.014943,-0.005684,0.004646,-0.016204,-0.005918,0.007872,0.004773,-0.000652,-0.003547,-0.003897,-0.000875,0.001373,-0.008656,-0.004314,0.001325,0.001676 Merovingian:KOS010,0.118376,0.139128,0.044877,0.027132,0.047393,0.020638,0.013161,0.010384,0.013908,0.027518,-0.007795,0.016335,-0.010258,-0.017065,0.016829,0.011138,0.019949,-0.015203,0.008799,0.012631,0.009358,0.001237,-0.011092,-0.000361,0.006347 Merovingian:KOS011,0.124067,0.137096,0.058077,0.036822,0.049548,0.027889,0.020681,-0.001154,0.013294,0.011481,-0.014615,0.001349,-0.009366,-0.018304,0.009636,0.013126,0.000522,-0.000253,0.004148,0.006878,-0.002246,0.006925,0.001356,0.017834,0.000359 Merovingian:KOS012,0.1161,0.11577,0.056568,0.048773,0.053548,0.018128,-0.001175,0.018922,-0.003886,0.000547,-0.008607,0.008692,0.001635,-0.005918,0.010179,0.020154,0.008214,-0.002154,-0.00817,0.009505,0.009234,0.005688,0.003204,-0.000241,0.010897 Merovingian:KOS013,0.12862,0.137096,0.060339,0.034884,0.034776,0.017291,0.004465,0.002538,0.004295,0.000729,-0.002436,0.01109,-0.008028,-0.00812,0.020765,0.006762,-0.007823,-0.002534,0.007668,0.001626,0.009234,-0.006925,-0.003081,0.013496,-0.012095 Merovingian:KOS015,0.122929,0.137096,0.060716,0.036499,0.035083,0.007251,-0.002585,0.001154,0.005522,0.011116,0.006983,0.007943,-0.01219,-0.006881,0.012622,-0.011933,-0.017732,0.000887,0.010182,0.003252,0.010232,0.002844,0.005546,0.015544,0.011256 Merovingian:KOS016,0.129758,0.13405,0.049026,0.035853,0.043393,0.017849,0,0.008307,0.005522,-0.001276,-0.009094,-0.001499,-0.005203,-0.007707,0.015744,0.009281,-0.021383,0.006714,0.007919,0.008629,0.004492,0.005564,0.006409,0.004699,0.000838 Merovingian:KOS017,0.126344,0.136081,0.053928,0.037791,0.038776,0.018407,-0.00094,0.006231,0.008181,-0.001093,-0.002436,0.000599,-0.010258,-0.007294,0.015201,0.001989,-0.015385,0.00114,0.005908,0.009505,0.009858,0.010881,0.002958,0.006748,0.002754 Merovingian:KOS022,0.125205,0.129988,0.057322,0.023256,0.040623,0.011713,0.01081,0.018692,-0.00225,0.008747,-0.003085,0.017385,-0.018583,-0.020506,0.02823,0.001989,-0.005607,-0.004434,-0.001006,0.004627,0.021462,-0.008161,-0.006655,0.012532,0.003113 Merovingian:KOS024,0.127482,0.141159,0.06675,0.043605,0.03416,0.019522,0.00047,0.006692,0.01309,0.012757,0.003735,0.001948,-0.019029,-0.02546,0.02538,0.009812,-0.013169,-0.006714,0.013198,0.004752,-0.004243,0.015828,0.006779,0.006507,-0.005389 Merovingian:KOS028,0.113823,0.13405,0.065242,0.046512,0.048009,0.019243,0,0.009923,0.007363,0.002734,-0.007307,0.004346,-0.009217,-0.004266,0.018187,0.007027,-0.002477,0.001394,-0.001006,0.001376,0.008859,0.000247,0.003697,0.013375,0.003353 Merovingian:KOS032,0.118376,0.142174,0.041483,0.028424,0.038469,0.004183,-0.008225,0.004384,0.00859,0.006925,-0.01088,0.007643,-0.009366,-0.012937,0.002307,-0.004906,-0.002738,0.002407,0,-0.003502,0.002371,0.006925,0.006286,-0.000241,-0.00479 Merovingian:KOS033,0.114961,0.127957,0.057322,0.037468,0.042777,0.020359,0.00987,-0.002769,0.015339,0.006378,-0.000812,-0.001649,-0.011596,-0.012799,0.016965,0.017369,0.019688,-0.007475,0.000126,0.00025,0.015223,-0.006554,0.001232,0.011447,0.005389 Merovingian:KOS034,0.120652,0.135065,0.058077,0.044897,0.041546,0.010598,0.00564,0.003461,0.014317,0.005467,-0.009906,0.001649,-0.008474,-0.012248,0.016558,0.006629,-0.004563,0.003421,-0.003017,0.002001,0.01148,0.001484,-0.002588,0.006627,-0.005987 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Merovingian:OLV050,0.133173,0.139128,0.054305,0.027778,0.035699,0.008646,0.011751,0.004615,0.007158,0.011116,0.000325,0.004046,-0.007136,-0.006331,0.010315,0.019623,0.011083,0.004687,0.001131,0.006128,0.009608,0.006183,0.002958,0.008555,-0.000838 Merovingian:OLV051,0.126344,0.140143,0.048649,0.028424,0.0397,0.015618,0.006345,0.005769,0.009204,0.016948,-0.003248,0.004796,-0.000297,-0.008395,0.015472,0.005304,-0.000782,0.011529,0.002388,0.003877,-0.004991,0.004081,0.002218,-0.000602,-0.000239 Merovingian:OLV054,0.126344,0.139128,0.055814,0.030685,0.040623,0.002789,-0.00282,-0.000923,0.003068,-0.005467,-0.003735,0.005695,-0.010406,-0.007019,0.014251,0.013657,0.003912,-0.003801,-0.006411,0.003502,0.002121,0.000989,-0.000493,0.011688,-0.003473 Merovingian:OLV055,0.114961,0.140143,0.063733,0.038437,0.0437,0.019522,-0.00564,0.003923,0.01084,0.015126,0.000974,0.012289,-0.015163,-0.007844,0.020629,-0.001458,-0.011343,0.010008,0.004022,0.005753,0.00262,0.000247,-0.006655,0.007832,-0.007305 Merovingian:OLV056,0.12862,0.152329,0.049403,0.024225,0.0437,0.013945,0.001645,-0.004615,0.008385,0.021322,-0.013803,0.002548,-0.013974,-0.002202,0.01045,-0.005038,-0.014994,0.004434,0.006285,-0.010755,-0.009982,-0.017435,-0.003081,0.016508,-0.013651 Merovingian:OLV058,0.119514,0.139128,0.047894,0.029393,0.042469,0.013945,0.000705,0.006923,0.013703,0.006925,0.000812,0.007943,-0.012042,-0.01156,0.01045,0.005038,-0.001956,-0.002534,-0.005531,0.007253,0.000624,0.000247,-0.003821,0.012652,-0.010897 Merovingian:OLV065,0.110408,0.127957,0.056945,0.020026,0.042777,0.007809,-0.001645,0.015461,0.006954,0.035354,-0.004872,-0.001199,-0.018285,-0.013487,0.007193,-0.001326,-0.003781,0.00228,0.009176,-0.003502,-0.011605,0.020279,-0.006532,0.010483,-0.006826 Merovingian:OLV067,0.118376,0.13405,0.0445,0.02261,0.036314,0.009761,0.00094,0.006692,0.004909,0.016401,-0.004547,0.005995,-0.00773,-0.013349,0.013843,0.018032,0.0103,-0.001647,-0.004399,-0.004752,0.010731,0.00272,0.000123,0.007591,0.008143 Merovingian:OLV071,0.127482,0.136081,0.059585,0.04845,0.037545,0.006414,0.007755,0.006923,0.005931,0.00656,0.001137,0.007943,-0.011596,-0.003028,0.011401,0.000928,-0.008475,0.001647,0.003771,0.009379,0.006738,0.003462,0.008997,0.005904,0.000599 Merovingian:OLV072,0.121791,0.13405,0.058831,0.01938,0.030775,0.014781,0.015041,0.008538,0.014112,0.016219,-0.007632,0.017085,-0.003865,-0.013487,0.005836,0.005967,-0.01343,-0.011275,0.00817,0.003126,0.013975,0.005935,0.006532,0.002651,0.001317 Merovingian:WPK001A,0.127482,0.13405,0.076178,0.047158,0.045239,0.022869,-0.011281,0.01223,0.007567,0.004191,-0.011854,-0.005695,-0.014123,-0.01445,0.017779,0.005171,0.002347,-0.002407,0.004022,0.006503,0.004866,0.000742,0.001232,0.002771,-0.007784 Merovingian:WPK002,0.135449,0.138112,0.058831,0.041344,0.036007,0.018686,0.010575,0.011307,0.005727,0.001093,-0.008769,0.008992,-0.011744,-0.014038,0.02158,-0.000796,-0.006128,0.004054,0.011061,0.003001,0.009982,0.001855,0.003944,0.000361,0.002754 Merovingian:WPK005,0.125205,0.140143,0.062602,0.040052,0.043393,0.01255,0,0.004615,0.009817,0.002187,-0.013316,0.003147,-0.006541,-0.009909,0.017915,0.011535,0.007041,0.003801,0.002137,0.000375,0.007487,0.012365,0.004067,0.008555,0.006227 Merovingian:WPK006,0.122929,0.13405,0.052797,0.04199,0.037238,0.016176,0.002585,0.005077,0.003068,0.004556,-0.00747,-0.00045,-0.010258,-0.016515,0.015879,0.012331,0.012908,0.005448,0.003142,0.01013,-0.001872,0.005317,0.008011,0.014098,-0.002036

 

References